Parajulid milliped studies V. The genera Pseudojulus Bollman and Arvechambus Causey (Parajulinae: Aniulini)

Rowland M. Shelley

Abstract


The parajulid milliped genus Pseudojulus Bollman comprises four species: P. obtectus (Bollman), P.paynei (Hoffman), n. comb., and P. carolinensis and P. coastalis, new species; Arvechambus Causey comprises two species, A. hummi and A. weemsi, both by Causey. Georgiulus Hoffman is placed in synonymy under Pseudojulus; G. hubrichti Hoffman is placed under P.paynei; and A. australis Causey is placed under A. hummi. The genera are sympatric in north Florida and southern Georgia, but Pseudojulus extends northward to coastal South Carolina and south central North Carolina, and westward to Alabama west of Mobile Bay. Both genera belong to the Aniulini and possess unique features; in Pseudojulus the anterior gonopod coxae are fused into a "shelf" on the dorsal surface of the complex that extends ventrad along the caudal margin and possesses a pair of posterior median syncoxal lobes or laminas of varying lengths and configurations. Arvechambus exhibits a suite of apomorphies and is sister to the rest of the tribe collectively: the 8th sternum possesses elevated lateral lobes that overhang the sides of the gonopodal aperture; the 7th pleurotergite possesses lobes that also overlie the sides of the aperture; the anterior gonopods lack lateral syncoxal processes, the coxal lobes, much larger than in other tribal genera, arise laterad and obscure part of the telopodite in anterior view, and the telopodite is elongate rather than clavate; and the 2nd pleurotergite in females possesses lobes that overhang and effectively close the cyphopodal aperture.

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