0263. Phylogeny and systematics of the giant rhinoceros beetles (Scarabaeidae: Dynastini)

J. Mark Rowland, Kelly B. Miller

Abstract


This report contains the first inclusive phylogenetic analysis and taxonomic structure for the genera presently included within the scarabaeid tribe Dynastini. The study was based upon morphological, biogeographic and molecular data and yielded direct support for the recognition of three subtribes in the Dynastini: Dynastina MacLeay, New Status (Dynastes Kirby, Augosoma Burmeister, Megasoma Kirby, Golofa Hope), Xylotrupina Hope, New Status (Xylotrupes Hope, Allomyrina Arrow, Trypoxylus Minck, Xyloscaptes Prell) and Chalcosomina Rowland and Miller, New Subtribe (Chalcosoma Hope, Haploscapanes Arrow, Beckius Dechambre, Eupatorus Burmeister, Pachyoryctes Arrow). The results provide indirect but significant support for the origin of the tribe Dynastini to be of an age no later than the early Late Cretaceous. Molecular and morphological evidence suggests that Eupatorus as constituted in current taxonomic literature is not monophyletic, and taxonomic alternatives are discussed to address this deficiency. A plausible explanation is also provided for the perplexing use by F. W. Hope of the family-group names Xylotrupidae and Dynastidae.

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